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When you're doing this threading it's really more about the processor than the gfx card isn't it? Are these threads on the CPU or GPU?

CPU. The rendering code is already very optimized and this is about eliminating all overhead on the CPU side.

as each thread is doing a smaller subset of the work, you are probably getting more cache hits, are you also using thread affinity on your busiest threads to stop them context switching.

 

I got the culling time down to an insanely low amount, and it would actually be much slower if I split it up into multiple threads:

 

sorry I assumed that each task was on a thread, running independently. I was suggesting that if you had more active threads than cpus, you would experience contention for those cpus. You would see this in the performance stats, as context switches, which will cause the current context to be saved, and another loaded. If this happens a lot you are loosing useful cpu processing power.